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Non-African populations, appeared shortly after 100,000 years ago.

A genetic split between the Khoisan and other Africans occurred roughly 260,000 years ago, shortly after humankind's origins and around the time of the Florisbad individual.

Khoisan people then diverged into two genetically distinct populations around 200,000 years ago, the researchers calculate. doi:10.1016/20) coaxed DNA from the remains of 15 ancient sub-Saharan Africans, from a variety of geographic regions dating from about 500 to 8500 years old.

Kung-san 'bushmen' of the Kalahari, the Sandawe and Hadzabe of Tanzania and the Biaka and Mbuti 'pygmies' of the Congo Basin have much to teach us both genetically and culturally.

The study also found that West Africans (green) can trace their lineage back to a human ancestor that may have split off from other African populations even earlier than the Khoe-San.

A missing link with a Near Eastern herder population that returned to Africa.

(W-Aa) Perforated Nassarius kraussianus shells from Blombos Cave.

(Ab) Conus ebraeus shell bead (Conus 2) from southern Africa (Border Cave). Images reproduced, with permission, from (A-D, H, I) The Stone Age Institute; (E-G, J-P, Ac) from [35]; (Q) from [47]; and (R-Ab) from [35,47,48].

A consistent and powerful hypothesis about human emergence is that the complementary reproductive strategies of females and males led to evolutionary gender paradox in early human societies and hence cultural complexity based on sexual relationships driven to a considerable extent by female reproductive choice.

The males, to achieve reproductive success needed to compromise their competition to fit with the cooperative nature of the human group, centered on the family and gathering and social relationships with the females.

Adaptions were found for resisting UV radiation (e.g.

in the Kalahari, and for bitter tastes providing better capacity to tell poisonous plants. (A-D) Bifacial foliates from northern Africa (A, Mugharet el Aliya; B-D, Adrar Bous).

Selection among males reinforces not just the traditional hunting prowess and toughness ('he-man') but diverse social skills ('domestic bliss') - "a mosaic of qualities that reflect the necessities of compromise ...

good with the children, relaxed, eloquent, knowledgeable".